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bioRxiv (Cold Spring Harbor Laboratory)

244 papers in the library · 999 citations · publishing 2015-2026

Papers

Experienced meditators exhibit no differences to demographically-matched controls in theta phase synchronisation, P200, or P300 during an auditory oddball task

bioRxiv (Cold Spring Harbor Laboratory) April 14, 2019 J.R. Payne, Oliver Baell, Harry Geddes et al. 6 citations preprint

Long-term meditation practice may not produce lasting changes in certain brain markers of attention. Experienced mindfulness meditators and non-meditators showed no differences in behavioral performance or in neural markers such as theta phase synchronisation, P200, and P300 during a dichotic oddball listening task when neither group meditated before or during the session. Bayes Factor analysis found evidence against group differences for P200 and P300. The findings suggest that increased activity in these attention-related neural markers does not reflect stable, trait-like changes in a community sample of mindfulness meditators, adding nuance to understanding how much meditation is needed to alter specific brain processes.

Striking Long Term Beneficial Effects of Single Dose Psilocybin and Psychedelic Mushroom Extract in the SAPAP3 Rodent Model of OCD-Like Excessive Self-Grooming

bioRxiv (Cold Spring Harbor Laboratory) June 29, 2024 Michal Brownstien, Michal Lazar, Alexander Botvinnik et al. 5 citations preprint

In mice with a genetic deletion that causes excessive self-grooming and anxiety—behaviors resembling aspects of obsessive-compulsive disorder (OCD)—a single dose of psilocybin or psychedelic mushroom extract reduced self-grooming over 21 days, while vehicle-treated mice showed a 118.7% increase. Psilocybin and the extract both decreased self-grooming by about 15–19%, and improvements in secondary measures like twitches and anxiety were also significant. In responsive mice, benefits lasted up to 7 weeks. The extract was superior for alleviating head-body twitches and anxiety. These results support clinical trials of psilocybin for OCD.

LSD flattens the hierarchy of directed information flow in fast whole-brain dynamics

bioRxiv (Cold Spring Harbor Laboratory) April 28, 2024 Kenneth Shinozuka, Prejaas K.b. Tewarie, Andrea I. Luppi et al. 5 citations preprint

LSD weakens the brain's directed connectivity hierarchy by increasing the balance between senders and receivers of neural signals. This finding supports the REBUS theory, which proposes that psychedelics flatten the hierarchy of information flow in the brain. Analyzing magnetoencephalography data from 16 healthy participants given 75 micrograms of intravenous LSD, the study found that LSD diminishes the asymmetry of directed connectivity averaged over time. Machine learning classifiers distinguished LSD from placebo more accurately when trained on hierarchy metrics than on traditional functional connectivity measures.

Sleep-like state during wakefulness induced by psychedelic 5-MeO-DMT in mice

bioRxiv (Cold Spring Harbor Laboratory) December 11, 2022 Benjamin J. B. Bréant, José Prius-Mengual, David M. Bannerman et al. 5 citations preprint

Psychedelics like 5-MeO-DMT induce a dissociated state of arousal that combines features of waking and sleep. In freely moving adult male mice, the drug produced sleep-like slow waves in the cortex alongside marked pupil dilation, even while animals were awake and moving. REM sleep was strongly suppressed, similar to the effect of conventional antidepressants. This mixed brain state may explain psychedelic effects such as dream-like hallucinations and reopening of the critical period for plasticity.

5-MeO-DMT modifies innate behaviors and promotes structural neural plasticity in mice

bioRxiv (Cold Spring Harbor Laboratory) November 3, 2022 Sarah J. Jefferson, Ian Gregg, Mark Dibbs et al. 5 citations preprint

The short-acting psychedelic 5-MeO-DMT increases head-twitch response in mice in a dose-dependent manner, with a shorter duration than psilocybin. It strongly suppresses social ultrasonic vocalizations during mating behavior and produces long-lasting increases in dendritic spine density in the medial frontal cortex by elevating the rate of spine formation, but unlike psilocybin, it does not affect spine size. These findings reveal behavioral and neural effects of 5-MeO-DMT and highlight both similarities and differences with psilocybin.

Role of 5-HT2A, 5-HT2C, 5-HT1A and TAAR1 receptors in the head twitch response induced by 5-hydroxytryptophan and psilocybin: Translational implications

bioRxiv (Cold Spring Harbor Laboratory) July 23, 2022 Orr Shahar, Alexander Botvinnik, Noam Esh-Zuntz et al. 5 citations preprint

Psilocybin and the serotonin precursor 5-HTP both cause a characteristic head twitch response in mice, a behavior linked to the human psychedelic experience. The head twitch response depends primarily on the 5-HT2A receptor, as blocking this receptor with volanserin reduced the response. Activating the 5-HT1A receptor also reduced head twitching. In contrast, blocking the 5-HT2C receptor had a bimodal effect, enhancing the response at lower doses but reducing it at higher doses. Blocking the trace amine associated receptor 1 (TAAR1) reduced head twitching caused by 5-HTP but not by psilocybin, indicating a differential role for this receptor. These findings identify multiple receptors that could be targeted to modulate the effects of psychedelic compounds in therapeutic settings.

Mapping Pharmacologically-induced Functional Reorganisation onto the Brain’s Neurotransmitter Landscape

bioRxiv (Cold Spring Harbor Laboratory) July 13, 2022 Andrea I. Luppi, Justine Y. Hansen, R. Adapa et al. 5 citations preprint

Psychoactive drugs reshape brain function by engaging multiple neurotransmitter systems simultaneously. By mapping the distribution of 19 neurotransmitter receptors and transporters (via PET) and the connectivity changes caused by 10 drugs (anesthetics, psychedelics, and stimulants), the study shows that drug effects are organized along hierarchical gradients of brain structure and function. Additionally, brain regions susceptible to drug-induced changes are also vulnerable to structural alterations from brain disorders. These findings reveal systematic links between molecular neurochemistry and large-scale functional reorganization.

Gamma band alterations and REM-like traits underpin the acute effect of the atypical psychedelic ibogaine

bioRxiv (Cold Spring Harbor Laboratory) June 29, 2020 Joaquín González, Matías Cavelli, Santiago Castro-Zaballa et al. 5 citations preprint

Ibogaine, a psychedelic alkaloid with anti-addictive properties, produces a waking state that shares brain-wave traits with REM sleep. In rats, ibogaine increased gamma oscillation power in the brain but made those oscillations less coherent and less complex than normal waking levels. This pattern mirrors REM sleep features within the gamma frequency band, providing biological evidence for the long-standing hypothesis that ibogaine induces a dream-like state while awake—a phenomenon called oneirogenesis. The findings offer an empirical basis for understanding how ibogaine's unique subjective effects may contribute to its anti-addictive potential.

Nightmares share strong genetic risk with sleep and psychiatric disorders

bioRxiv (Cold Spring Harbor Laboratory) November 9, 2019 Hanna M. Ollila, Nasa Sinnott-Armstrong, Katri Kantojärvi et al. 5 citations preprint

Nightmares are vivid, disturbing dreams that wake the dreamer. Twin studies show nightmare frequency is 36–51% heritable. A genome-wide association study of 28,596 individuals from Finland and the United States identified genetic variants near MYOF and PTPRJ (a gene previously linked to sleep duration) that predispose to nightmares. Strong genetic correlations were found between nightmare frequency and neuroticism, post-traumatic stress disorder, major depressive disorder, daytime sleepiness, and insomnia. Mendelian randomization indicated that feelings of being fed-up, nervous, and miserable cause nightmares, with no evidence that nightmares cause psychological problems. The findings suggest nightmares arise from unique genetic risk factors and are substantially caused by underlying psychological and sleep problems.

Psilocybin exerts distinct effects on resting state networks associated with serotonin and dopamine in mice

bioRxiv (Cold Spring Harbor Laboratory) September 1, 2019 Joanes Grandjean, David Buehlmann, Michaela Buerge et al. 5 citations preprint

Psilocybin, a serotonin 2A receptor agonist, alters functional connectivity in the brain's default-mode network, which is involved in self-reference and disrupted in depression. In lightly-anesthetized mice, resting-state fMRI showed psilocybin reduced connectivity within the ventral striatum. Using gene expression maps and viral tracer projections, two distinct effects emerged: psilocybin increased connectivity between serotonin-associated networks and parts of the mouse default-mode network, thalamus, and midbrain, while decreasing connectivity within dopamine-associated striatal networks. These findings suggest that interactions between serotonin- and dopamine-regulated neural networks contribute to psilocybin's neural and psychological effects, and show how molecular and structural connectivity data can clarify pharmaco-fMRI results.

A Single Dose Of Ayahuasca Modulates Salivary Cortisol In Treatment-Resistant Depression

bioRxiv (Cold Spring Harbor Laboratory) January 31, 2018 Ana Cecília de Menezes Galvão, Raíssa Nóbrega de Almeida, Erick Allan Dos Santos Silva et al. 5 citations preprint

In treatment-resistant depression, a single dose of ayahuasca normalizes the blunted awakening salivary cortisol response that is characteristic of the disorder. Patients with major depression showed hypocortisolemia and a diminished cortisol awakening response compared with healthy controls at baseline. During the dosing session, both patients and controls who ingested ayahuasca had a large increase in salivary cortisol relative to placebo groups. Forty-eight hours after ayahuasca, the awakening cortisol response in treated patients became similar to that of controls, an effect not seen with placebo. No changes in plasma cortisol occurred 48 hours after either ayahuasca or placebo. The modulation of salivary cortisol may contribute to ayahuasca's rapid antidepressant effects.

Lasting effect of psilocybin on sociability can be blocked by DNA methyltransferase inhibition

bioRxiv (Cold Spring Harbor Laboratory) March 11, 2025 Chenchen Song, Tinya Chang, Tobias Buchborn et al. 4 citations preprint

A single dose of the psychedelic psilocybin lastingly improves social behavior in a mouse model of autism (Cntnap2-knockout mice). This effect is blocked by inhibiting DNA methyltransferase I, suggesting an epigenetic mechanism involving DNA methylation underlies psilocybin's long-term influence on social function.

Long-term effects of psilocybin on dynamic and effectivity connectivity of fronto-striatal-thalamic circuits

bioRxiv (Cold Spring Harbor Laboratory) November 7, 2024 Lorenzo Pasquini, Jakub Vohryzek, Anira Escrichs et al. 4 citations preprint

Psilocybin induces fast and sustained improvements in mental well-being, yet its long-term mechanisms are not fully understood. Four weeks after a full dose, fronto-striatal-thalamic (FST) circuitry—involved in goal-directed behavior and motivation—shows increased dynamic activity and flexibility in healthy volunteers. Computational modeling indicates that reduced structural constraints on functional dynamics cause this increased flexibility. Long-term changes include increased bottom-up and reduced top-down information flow, mediated by serotonergic (5-HT2A) and dopaminergic (D2) receptor systems. This functional re-organization of FST circuits may represent a common mechanism underlying clinical improvements across neuropsychiatric disorders such as substance abuse, major depression, and anorexia.

The serotonin 1B receptor is required for some of the behavioral effects of psilocybin in mice

bioRxiv (Cold Spring Harbor Laboratory) October 21, 2024 Sixtine Fleury, Katherine M. Nautiyal 4 citations preprint

Psilocybin's persisting clinical effects are commonly attributed to activation of the serotonin 2A receptor, but its active metabolite binds to many serotonin receptor subtypes, including the serotonin 1B receptor (5-HT1BR). In mice, 5-HT1BR expression influenced brain-wide activity after psilocybin administration, measured by differences in c-Fos patterns across regions involved in emotional processing and cognitive function, including the amygdala and prefrontal cortex. 5-HT1BR mediated some acute and persisting behavioral effects: mice lacking 5-HT1BRs showed attenuated hypolocomotion to psilocybin, and both transgenic and pharmacological loss-of-function models indicated 5-HT1B involvement in decreased anhedonia and reduced anxiety-like behavior. The research implicates 5-HT1BR as a mediator of psilocybin's behavioral and neural effects in mice.

A dual-receptor model of serotonergic psychedelics

bioRxiv (Cold Spring Harbor Laboratory) April 15, 2024 Arthur Juliani, Veronica Chelu, Laura Graesser et al. 4 citations preprint

Serotonergic psychedelics show promise for treating mood and anxiety disorders, but their therapeutic mechanism remains debated. A popular theory holds that strong 5-HT2a receptor activation disrupts cortical dynamics, loosening rigid, maladaptive beliefs and making them open to revision. This work extends that perspective by developing a simple energy-based model of cortical dynamics rooted in predictive processing and neuromodulation. The model simulates hypothetical computational mechanisms for both 5-HT2a and 5-HT1a agonism, and its results account for several existing empirical observations about psychedelics' effects on cognition and affect. The model provides a theoretically grounded hypothesis for the clinical success of LSD, psilocybin, and DMT, and identifies biased 5-HT1a agonist psychedelics like 5-MeO-DMT as potentially fruitful for developing more effective and tolerable psychotherapeutic agents.

Cultivation, chemistry, and genome of Psilocybe zapotecorum

bioRxiv (Cold Spring Harbor Laboratory) November 2, 2023 Jordan Taylor Jacobs, Alan Rockefeller, Harte Singer et al. 4 citations preprint

*Psilocybe zapotecorum*, a mushroom with historical indigenous use, contains a potent psilocybin concentration of 17.9 mg/g, ranging from 10.6-25.7 mg/g across seven samples. Its full genome, revealing the psilocybin gene cluster, offers new biological insights. Combining Fungal Biology and Applications with Chemical synthesis and alkaloids, the work profiles its complex tryptamine chemistry, including psilocin and baeocystin. A total tryptamine concentration reached 22.5 mg/g. These findings advance Psychedelics and Drug Studies by illuminating the diversity of naturally occurring psilocybin sources.

Dose-response relationships of LSD-induced subjective experiences in humans

bioRxiv (Cold Spring Harbor Laboratory) November 7, 2022 Tim Hirschfeld, Johanna Prugger, Tomislav Majić et al. 4 citations preprint

Lysergic acid diethylamide (LSD) produces a sigmoid-like increase in altered states of consciousness, with effects plateauing around 100 micrograms. The strongest changes involve perception and illusory imagination, followed by positively experienced ego-dissolution, while anxiety and dread of ego dissolution show only small effects. Considerable variability in responses highlights the importance of non-pharmacological factors. These dose-response relationships can serve as references for future research on LSD.

Ritualistic use of ayahuasca enhances a shared functional connectome identity with others

bioRxiv (Cold Spring Harbor Laboratory) October 11, 2022 Pablo Mallaroni, Natasha L. Mason, Lilian Kloft et al. 4 citations preprint

Brain functional connectomes are unique and reliable identifiers of individuals, but it was unknown whether these 'fingerprints' persist during altered states of consciousness. Ayahuasca, a serotonergic psychedelic, disrupts functional connectivity. In a within-subject study using 7T fMRI, 21 members of the Santo Daime church were scanned after collective ayahuasca intake. Connectome fingerprinting revealed a shared functional space and a spatiotemporal reallocation of key edges. Differences in higher-order functional connectivity motifs predicted perceptual drug effects, showing that individualized connectivity markers can trace a subject's functional connectome across altered states.

Individual and combined effects of Cannabidiol (CBD) and Δ9-tetrahydrocannabinol (THC) on striato-cortical connectivity in the human brain

bioRxiv (Cold Spring Harbor Laboratory) November 21, 2020 Matthew B. Wall, Tom P. Freeman, Chandni Hindocha et al. 4 citations preprint

Cannabidiol (CBD) and Δ9-tetrahydrocannabinol (THC) are two major cannabis constituents with contrasting actions: THC is psychoactive and addiction-promoting, while CBD may have opposite effects. In two placebo-controlled, double-blind studies, inhaled THC (8 mg) strongly disrupted functional connectivity in associative and sensorimotor striatal networks, and this disruption was selectively alleviated in the limbic striatum when co-administered with CBD (10 mg). Oral CBD (600 mg) alone increased connectivity in the associative network but caused minor decreases in limbic and sensorimotor networks. The insula emerged as a key region affected by cannabinoid-induced connectivity changes, with implications for cannabis-related disorders and cannabinoid therapeutics.

Dopamine manipulations modulate paranoid social inferences in healthy people

bioRxiv (Cold Spring Harbor Laboratory) December 19, 2019 J.M. Barnby, Vaughan Bell, Quinton Deeley et al. 4 citations preprint

Dopamine transmission influences social attributions related to paranoia, but not the salience of paranormal or other beliefs. In a double-blind, placebo-controlled experiment, 27 healthy men received either L-DOPA (150 mg), haloperidol (3 mg), or placebo across three sessions. Haloperidol reduced attributions of harmful intent in a Dictator Game, while L-DOPA reduced such attributions only in fair conditions. Haloperidol unexpectedly increased attributions of self-interest for opponents' decisions. No changes occurred in belief salience for politics, religion, science, morality, or the paranormal. These results suggest dopamine selectively affects social inferences linked to paranoia, independent of mood or skepticism.

Reduced responsiveness of the reward system underlies tolerance to cannabis impairment in chronic users

bioRxiv (Cold Spring Harbor Laboratory) July 19, 2019 Natasha L. Mason, Eef L. Theunissen, Nadia R. P. W. Hutten et al. 4 citations preprint

Regular cannabis users develop tolerance to the drug's impairing and rewarding effects, but the underlying brain changes are unclear. In a double-blind, randomized, placebo-controlled crossover study, 12 occasional and 12 chronic cannabis users received acute doses of cannabis (300 μg/kg THC) and placebo, then underwent ultra-high field fMRI and magnetic resonance spectroscopy. In occasional users, cannabis caused decreased functional connectivity and increased striatal glutamate concentrations in reward circuitry, linked to greater subjective high and worse sustained attention. These changes were absent in chronic users, suggesting reduced responsiveness of reward circuitry to cannabis intoxication and a pharmacodynamic mechanism for tolerance development.

Acute antidepressant effect of ayahuasca in juvenile non-human primate model of depression

bioRxiv (Cold Spring Harbor Laboratory) January 25, 2018 Flávia Santos Da Silva, Erick Allan Dos Santos Silva, Geovan Menezes de Sousa et al. 4 citations preprint

In a juvenile model of depression using common marmosets, a single dose of ayahuasca reversed depressive-like symptoms within 24 hours, including recovery of cortisol levels, reduced stereotypic scratching in males, increased feeding, and restored body weight in both sexes. The effects lasted 14 days. The study suggests ayahuasca produces faster and more durable antidepressant effects than the tricyclic antidepressant nortriptyline, supporting its potential as a treatment for early-age depression.

Integrated phenomenology and brain connectivity demonstrate changes in nonlinear processing in jhana advanced meditation

bioRxiv (Cold Spring Harbor Laboratory) Ruby M. Potash, Sean D. Van Mil, Mar Estarellas et al. 4 citations preprint

During a deep meditative state called jhana, the brain's non-oscillatory, nonlinear neural activity—rather than oscillatory synchrony—best distinguishes the state from ordinary waking consciousness. In a single highly experienced meditator (over 20,000 hours of practice) studied across 29 sessions, EEG recordings showed that combining subjective ratings of attention with a nonlinear connectivity metric improved the ability to decode the meditative state compared to using neural measures alone. Deeper jhana states were marked by a balance between feedback and feedforward neural processes, indicating an equalization of internally and externally directed information processing. These findings suggest that refined conscious states involve distinct large-scale neural dynamics not captured by traditional oscillatory measures.

Psilocybin Prolongs the Neurovascular Coupling Response in Mouse Visual Cortex

bioRxiv (Cold Spring Harbor Laboratory) July 31, 2025 Rick Zirkel, Matthew Isaacson, Clara Liao et al. 3 citations preprint

Psilocybin prolongs increases in visual stimulus-evoked capillary blood flow in the mouse visual cortex without altering stimulus-evoked neural activity. This effect was reduced by pretreatment with a 5-HT2A receptor antagonist. Multi-modal widefield imaging confirmed extended vascular responses in surface vessels with no observed effect on population neural response. Computational simulations showed that prolonged neurovascular coupling responses can produce spurious increases in BOLD-based measures of functional connectivity. These findings demonstrate that psilocybin broadens neurovascular responses in the brain, highlighting the need to account for these effects when interpreting human neuroimaging data of psychedelic drug action.

PsiConnect: A Multimodal Neuroimaging Study of Psilocybin-Induced Changes in Brain and Behaviour

bioRxiv (Cold Spring Harbor Laboratory) April 14, 2025 Leonardo Novelli, Devon Stoliker, Tamrin Barta et al. 3 citations preprint

PsiConnect is a large-scale neuroimaging study that examined brain activity in 62 participants before and after a 19 mg dose of psilocybin using functional, structural, and diffusion-weighted MRI combined with EEG. The design included resting-state scans and three naturalistic conditions: guided meditation, music listening, and movie watching. Half of the participants completed an 8-week meditation training program, allowing exploration of interactions among meditation, psilocybin, and brain function. Multi-echo fMRI improved signal-to-noise ratio and reduced artifacts. Behavioral and self-report measures captured acute and longitudinal effects, with follow-ups extending to one year. The data is curated according to open science principles.