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The psychedelic DOI decreases the modularity of the cortical network by increasing cortico-cortical and thalamo-cortical functional influence

Roman Doronin, Eduard Stroukov, Georg B. Keller

bioRxiv (Cold Spring Harbor Laboratory) July 24, 2026 preprint DOI: 10.64898/2026.07.20.739656 (opens in new tab)

Study at a glance

AI-extracted from the abstract
Characteristics Experimental study
Population Awake mice
Intervention DOI
Key findings DOI acutely decreased cortical network modularity by increasing correlations between anterolateral and posteromedial cortical domains, dependent on 5-HT2A receptors, and increased thalamo-cortical and long-range cortico-cortical functional influence onto the anterolateral domain.

Abstract

SUMMARY Psychedelics are powerful modulators of conscious perception, hypothesized to exert their effects by reorganizing large-scale brain functional connectivity. The circuit mechanisms underlying this reorganization remain unknown. Here, using widefield calcium imaging in awake mice, we found that the psychedelic DOI, in a 5-HT2A-dependent manner, acutely decreased the modularity of the cortical network by increasing correlations between anterolateral (AL) and posteromedial (PM) cortical domains. Functional influence measurements revealed that DOI increased both thalamo-cortical and long-range cortico-cortical influence onto the AL domain. We speculate that these increases in functional influence are driving the DOI-induced increase in the coupling between cortical domains. Our results provide a mouse correlate of the network reorganization seen under psychedelics in human neuroimaging and identify thalamo-cortical and cortico-cortical changes as candidate circuit mechanisms for this reorganization.